A flow control element (20) is provided with a resilient sealing flange (54) biased to seal in-venting passages (36) in a housing (22) around the element (20). The element (20) also includes a self-sealing, pressure-openable, dispensing valve (70). The element (20) and housing (22) can be combined to form a dispensing structure that can function to discharge a fluent material product out of the dispensing structure through the dispensing valve (70) while simultaneously venting exterior ambient air (or other exterior ambient environment fluent material) into the dispensing structure.