The invention relates to the technical field of harvesters, in particular to a stone discharging device and a harvester. The stone discharging device comprises a stone discharging door, a door openingand closing mechanism and a triggering mechanism; the door opening and closing mechanism is used for making the stone discharging door opened or closed; the triggering mechanism is installed on the stone discharging door and used for making the door opening and closing mechanism act to open the stone discharging door. The harvester comprises a threshing device and the stone discharging device, and the stone discharging door of the stone discharging device is located on a crop conveying route between an extruding roll and a cutter of the threshing device. The stone discharging device can avoidthat stones influence the threshing device and a separating device, in this way, the service life of the harvester is prolonged, and the effects of threshing and separating are improved.