A network communication system includes a network node that is configured to create and terminate sessions between talkers and listeners connected to different Layer 2 networks. The talkers and listeners may be configured to reserve network paths and communicate data streams in accordance with a Layer 2 reservation protocol, such as Stream Reservation Protocol. The talkers and listeners in the different Layer 2 networks may reserve network paths and communicate data streams through a network node configured to manage sessions using a Layer 3 session protocol, such as Session Initiation Protocol. The network node may reserve the network paths and enforce quality of service in accordance with the Layer 2 reservation protocol so that the talkers and listeners in the different Layer 2 networks may communicate data streams to each other.