An actuating system is provided for a dispenser cartridge that can be mounted on a frame and that has a reciprocatable product-dispensing hollow stem that (A) is biased to an extended position in which the cartridge is unactuated, and (B) is moveable from the extended position to a depressed position in which the cartridge is actuated for discharging the fluent substance through the stem. The actuating system includes a plunger on the frame, and a discharge conduit located adjacent the frame to accommodate relative movement between the discharge conduit and the frame. The discharge conduit includes (A) ml inlet opening communicating with the dispenser cartridge hollow stem, (B) ml outlet opening from which the fluent substance can be dispensed, (C) a passageway between the inlet opening and outlet opening, and (D) an intermediate opening that is in communication with the passageway between the inlet opening and the outlet opening.