Users may dynamically specify a “posing root” node in an animation hierarchy that is different than the model root node used to define the animation hierarchy. When a posing root node is specified, users specify the pose, including translations and rotations, of other nodes relative to the posing root node, rather than the model root node. Poses of nodes may be specified using animation variable values relative to the posing root node. Animation variable values specified relative to the posing root node are dynamically converted to equivalent animation variable values relative to the model root node, which then may be used to pose an associated model. Animation data may be presented to users relative to the current posing root node. If a posing root node is changed to a different location, the animation data is converted so that it is expressed relative to the new posing root node.