In an audio processing system (300), a filtering section (350, 400): receives subband signals (410, 420, 430) corresponding to audio content of a reference signal (301) in respective frequency subbands; receives subband signals (411, 421, 431) corresponding to audio content of a response signal (304) in the respective subbands; and forms filtered inband references (412, 422, 432) by applying respective filters (413, 423, 433) to the subband signals of the reference signal. For a frequency subband: filtered crossband references (424, 425) are formed by multiplying, by scalar factors (426, 427), filtered inband references of other subbands; a composite filtered reference (428) is formed by summing the filtered inband reference of the subband (422) and the filtered crossband references; a residual signal (429) is computed as a difference between the composite filtered reference and the subband signal of the response signal corresponding to the subband; and the scalar factors and the filter applied to the subband signal of the reference signal corresponding to the subband are adjusted based on the residual signal.