The present application discloses a touch substrate. The touch substrate includes a base substrate, and a first mesh electrode layer having a plurality of first mesh electrodes on the base substrate. The plurality of first mesh electrodes are arranged substantially along a first direction. Each of the plurality of first mesh electrodes extends substantially along a second direction. Each of the plurality of first mesh electrodes includes a plurality of first portions having a zig zag contour, a midline of each of the plurality of first portions being a zig zag line.