A display substrate includes a base substrate and a unitary indivisible pixel electrode. The base substrate includes a gate line, a data line crossing the gate line and a thin-film transistor electrically connected to the gate line and the data line formed thereon. A pixel area is defined on the base substrate. The unitary indivisible pixel electrode includes a plurality of branch electrodes formed on the pixel area and a connection electrode connecting the branch electrodes to each other. Each of the branch electrodes includes a bent portion and the bent portions of adjacent branch electrodes are spaced apart from each other.