A fuel cell stack includes a plurality of membrane electrode assemblies, each of the membrane electrode assemblies having an electrolyte membrane; an anode on a first side of the electrolyte membrane; and a cathode on a second side of the electrolyte membrane opposite to the first side, wherein the anode and the cathode each comprise a gas diffusion layer divided into at least two areas such that a first area and a second area have different area densities; and a separator between adjacent membrane electrode assemblies.