A banknote accommodating device includes a box body, a box door and a banknote inlet. A movable guide plate in the box body is arranged below the banknote inlet through a rotating shaft, and has a guide surface facing the banknote inlet, and an included angle between the guide surface and a vertical direction ranges from 0 degree to 20 degrees, an elastic element is provided on a back surface of the guide surface, and provides a thrust to rotate the movable guide plate counterclockwise. The box door is provided with a guide rib which forms an included angle with the vertical direction, and a stacking surface is provided at the bottom of the box body and forms an included angle with a horizontal direction, and a space for stacking banknotes vertically is formed between the guide surface, the guide rib and the stacking surface.