A display substrate and a display device. The display substrate includes a base and a shift register unit provided on the base; the shift register unit includes a first first voltage line, an on-off control circuit, a first energy storage circuit and a fourth node control circuit; the on-off control circuit comprises a first transistor, and the first energy storage circuit includes a first capacitor; at least part of the first first voltage line extends in the a first direction; the fourth node control circuit includes a second transistor; the first capacitor, the first transistor, the first first voltage line and the second transistor are arranged in a second direction; the second transistor, the first first voltage line, the first transistor and the first capacitor are arranged in sequence along a direction close to the display area; the first direction intersects the second direction.