A wheel securing adapter is provided in a bicycle fork to attach a skewer of a wheel securing axle thereto. The wheel securing adapter is disposed in an axle mounting opening of the fork. The wheel securing adapter has an internally threaded bore and an external surface configured and arranged to cooperate with the axle mounting opening to prevent at least one of relative axial movement and relative rotational movement of the wheel securing adapter within the axle mounting opening. The fork includes a pair of fork legs having upper ends coupled to the fork stem and lower ends with axle mounting openings. Preferably, a cutout surrounds one of the axle mounting openings on an inner side surface that faces the other fork leg to form the mounting opening for the wheel securing adapter.