A liquid ejection substrate used to eject a liquid includes a substrate having an element row including a plurality of elements provided in a row, where each of the elements generates energy, drive circuits that drive and control the elements, a signal line that is shared by and connected to the drive circuits, the signal line being provided along the element row, and a heating portion generating heat used to heat the substrate, wherein the heating portion is provided so that the heating portion does not overlap the signal line with reference to a direction perpendicular to a face of the substrate.