An open roof construction for a vehicle includes a first closing element movably supported by a stationary part and is configured to move to an upward position with at least its front edge. A second closing element movably supported by the stationary part and is positioned rearward of the first closing element. The second closing element moves between a closed position in which it closes said roof opening and an at least partly open position. An air guide on a lower side of the first closing element guides and directs the air caught by said first closing element when in its upward position during driving of the vehicle to create a boundary layer of air which is situated between the outside and the inside of the vehicle at the position of the roof opening when said second closing element is in its at least partly open position.