Provided are (i) a touch panel substrate having improved uniformity in in-plane light transmittance and (ii) an electronic device including that touch panel substrate. First electrodes (11) each include a first protrusion (15), the first protrusions (15) each have an intersection at which two conductor lines cross each other, second electrodes (21) each include a second protrusion (25), the second protrusions (25) each have an intersection at which two conductor lines cross each other, and the plurality of first electrodes (11) and the plurality of second electrodes (21) are placed in such a manner that the plurality of first protrusions (15) and the plurality of second protrusions (25) engage alternately with each other in a plan view.