Provided are a planographic printing plate precursor which includes a hydrophilic support and an image recording layer containing an organic-inorganic hybrid particle (A) and an infrared-ray absorbent (B) on the hydrophilic support, in which the organic-inorganic hybrid particle (A) is an organic-inorganic hybrid particle in which an inorganic particle (a) containing a hydroxyl group and an organic component (b) are linked to each other through a linking group (c) having a urethane bond or a urea bond; a method of preparing a planographic printing plate using the planographic printing plate precursor; a planographic printing plate precursor which is capable of achieving both of printing durability and developability at high levels by using an organic-inorganic hybrid particle in which an inorganic particle (a) containing a hydroxyl group and an organic component (b) are linked to each other through a linking group (c) having a urethane bond or a urea bond, and the organic component (b) is linked to at least two inorganic particles (a) and the inorganic particle (a) is linked to at least two organic components (b) so that the organic components and the inorganic particles form a network structure; a method of preparing a planographic printing plate using the planographic printing plate precursor; and the organic-inorganic hybrid particle used for the planographic printing plate precursor and the method of preparing a planographic printing plate.