An air filter element includes (a) an open filter interior having a first closed end and a second open end opposite the first closed end; the second open end having an air flow aperture therethrough; (b) filter media provided in a pleated configuration and arranged around the open filter interior between the first closed end and the second open end; (c) a seal arrangement positioned at the open end of the filter element and includes an inner perimeter radial facing surface and an opposite, outer perimeter radial facing surface, wherein the inner perimeter radial facing surface includes a seal surface, and wherein one of the inner perimeter radial facing surface and the outer perimeter radial facing surface is a circular surface and the other of the inner perimeter radial facing surface and the outer perimeter radial facing surface is a non-circular surface.