A tiltable vehicle has a pair of front wheels coupled to a tiltable chassis by a tilt linkage, such that the pair of front wheels and the chassis are configured to tilt in unison with respect to a roll axis of the chassis. The tilt linkage includes a four-bar linkage having a pair of upper bar segments coupled to the chassis at spaced-apart respective inboard joints. In some examples, the inboard joints of the upper bar segments are each disposed outboard of a central chassis joint of a lower bar of the tilt linkage. An orientation sensor is configured to detect directional information regarding a net force vector applied to the chassis, and a tilt actuator operatively coupled to the chassis and configured to selectively tilt the chassis. A controller is configured to selectively control the tilt actuator based on the directional information from the orientation sensor.