IR-sensitive lithographic printing plate precursors provide a stable printout using a unique IR-sensitive image-recording layer. The IR radiation-sensitive layer includes: component (1) a free radical initiator composition; component (2) a free radically polymerizable composition; and component (3) a color-changing compound of Structure (I) having an indene structure in the conjugated chain between the aromatic terminal groups. A specific group, such as a halo group, is directly or indirectly attached to this indene structure. The infrared radiation-sensitive image-recording composition and layer also contains one or more borate ions. After infrared imaging, these precursors exhibit desirable fresh or initial printout and printout after dark storage. The precursors can be developed on-press.