Fuel cell systems (10) and related methods for limiting fuel cell slippage are provided. A stacked plurality of adjacent fuel cells (14) collectively forming a fuel cell stack (12). The fuel cells each include a pair of first and second plates (30, 30′, 30″; 32, 32′, 32″) at respective opposite ends thereof. A first fuel cell has a first plate (30, 30′, 30″) in engagement with a second plate (32, 32′, 32″) of a second fuel cell adjacent to the first fuel cell. A slip mitigation arrangement (50, 50′, 50″) between at least one of the pairs of the first and second fuel cells comprises first and second seats (62, 62′, 62″; 64, 64′, 64″) recessed in the engagement surfaces of the first and second conductive plates respectively, and a key member (60, 60′, 60″) having opposite ends seated in the first and the second recessed seats such that relative movement between the first and the second fuel cells is limited.