A head cartridge and a liquid ejection apparatus are provided in which cleaning performance of cleaning means is improved using a liquid absorbing force of a wiping member produced along with restoration of temporarily increased elastic displacement of the wiping member. An elastic displacement h (height of a projection) of a cleaning roller temporarily produced by the projection arranged at a position in the foreground of ink ejection nozzles in the cleaning direction of a nozzle surface is established to satisfy the following condition: h>(Vu/Vr)(L+n/2−φ/2), where the restoring speed of the elastic deformation of the cleaning roller is denoted as Vu; the moving speed of the cleaning roller is denoted as Vr; the movement distance of the cleaning roller from a restoring initiation point of the elastic deformation to the center of the liquid ejection nozzles is L; the contact width between the cleaning roller and the nozzle surface is n; and the diameter of the ink ejection nozzle is φ.